association	dataset	threshold value	standardized value
ZNF768	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	38.6114
TBXT	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	35.214
PPARG	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	47.8571
RELA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	21.6667
ASCL1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	77.6297
GLIS3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	34.6273
HAND2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	19.7468
ZNF669	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	32.02
GLI3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	47.1618
ELF1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	32.7088
MSANTD3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	20.4243
ZFP14	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	75.2349
ZNF784	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.5929
NR1I2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	23.3364
EWSR1-FLI1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	237.819
NKX2-3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	22.9039
THAP1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	57.816
ZEB1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.4204
PORCN	CM4AI KOLF21J CRISPRi Gene Perturbation Atlas	-1.0	-2.02534
PRDM1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	51.2416
DMRTA2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.4183
PAX9	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	37.6605
CREB3L1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	24.0994
TEF	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	26.7619
ARNT::HIF1A	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	14.2212
JDP2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	49.3314
MAFK	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	27.3325
ELK4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	42.6315
NKX2-4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	16.4828
RARA::RXRG	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.0785
EBF1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	57.357
SPIB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	62.3155
ZNF263	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	40.5383
ETV5::FOXO1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.7788
REL	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.1176
BCL11A	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	32.8097
ZNF682	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	84.4915
CTCF	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	230.223
ZNF75D	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	23.8544
ZNF528	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	41.6867
FLI1::DRGX	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	22.9395
MAX::MYC	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	20.3333
TEAD2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	39.8026
MAF	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	59.1406
ZNF175	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.4925
NRL	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	22.4164
DMRTA1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	24.384
FOSL2::JUND	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	45.9128
FOSL1::JUN	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	44.1374
STAT3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	24.0422
VEZF1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	37.3167
ZNF530	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	73.9697
ZNF416	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	33.4348
ZNF281	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	148.737
ZIC1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	58.9988
ZBTB6	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.9079
JUNB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	42.8397
NEUROD1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	29.6775
FOSL1::JUNB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	23.927
ZNF257	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	64.6767
SIX2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	33.6151
ATOH7	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.9861
EHF	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.2464
COPS4	CM4AI KOLF21J CRISPRi Gene Perturbation Atlas	-1.0	-2.15546
ZNF701	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	81.5943
SOX10	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	15.6117
RFX4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.3174
ELF3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.3534
OVOL2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.1486
MAFA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	51.8408
NFIL3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	18.3275
SNAI3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	32.4483
FOSB::JUNB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	38.5953
FOXO1::ELK3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.1555
IRF5	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	23.8602
TCF4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	35.8012
TP53	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	37.7459
HES7	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	55.9933
CTCFL	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	48.93
ELF2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	19.2385
PRDM9	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	164.867
ZBTB24	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	57.1232
ZNF740	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	140.397
ERF::FIGLA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	88.3564
ZNF547	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.2426
ZNF582	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	50.9352
NR1D1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	66.1418
GABPA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.8167
PLAG1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	110.0
JUND	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	41.1978
MTF1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	26.4257
NR2F1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	63.5542
KLF14	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	89.5212
ZNF157	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	33.3657
CREM	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	20.2964
CEBPG	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	27.3382
TCF7L1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	46.0867
ZNF324	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	38.7623
HEY1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	19.467
SNAI1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	22.2807
ZNF680	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	42.659
NFIC	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	73.6882
SCRT2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	40.2471
DMRTC2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	39.8868
ETV5::FOXI1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.5374
STAT1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	38.8843
TBX1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	18.0798
CLOCK	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	21.416
STAT1::STAT2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	69.6806
RREB1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	166.0
ZNF148	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	124.7
ZNF549	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	47.7487
FOS::JUNB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.9175
PKNOX1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	20.6173
TBX5	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	16.0773
TFEB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	16.975
PITX3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	14.5523
FOS::JUN	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.0349
ZNF675	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	37.631
MAF::NFE2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	52.8119
ESRRB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.2188
MGAT3	CM4AI KOLF21J CRISPRi Gene Perturbation Atlas	1.0	2.36072
MSC	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	27.7141
PMC12356745-12015_2025_10896_MOESM2_ESM.pdf-886-ENSG00000184481_FOXO4	Rummagene Transcription Factor Associations 2026	1.0	null
MYC	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.4947
NFIC::TLX1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	52.3125
ZNF320	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	179.569
REST	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	84.4159
FOSL2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	34.2757
PAX2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	34.1773
IKZF2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.5706
ZSCAN31	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	30.673
KLF5	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	108.837
ZNF214	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	23.2637
FOSL2::JUN	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.6197
HES5	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	30.1045
ZNF274	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	22.5283
PBX3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	48.9902
GRHL2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	23.0581
MAFF	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.3969
ERF::SREBF2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	19.1403
TFAP4::FLI1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	61.9333
CREB3L4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	43.0764
BACH1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	34.5831
FOXO1::FLI1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	32.2796
NEUROG2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	34.1
SCRT1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	34.2179
MXI1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.3262
TFAP4::ETV1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	39.7038
ZNF449	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	47.4104
ZNF816	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	50.8196
KLF4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	73.855
RFX5	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.71
ATF6	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	41.9737
FOS	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	49.51
PBX1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.5
OLIG3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	18.442
PAX1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	44.4556
OSR2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.414
MYF5	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	23.2281
NFE2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	23.9136
ATF3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	21.0821
FEZF2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	77.9359
NFIB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	83.0793
ZKSCAN3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	66.6619
SNAI2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	37.0382
TBX21	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	27.24
OSR1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	41.9367
TCF12	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	44.9541
MAX	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	19.6591
ZNF770	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	54.2402
ESRRA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.3885
PMC12356745-12015_2025_10896_MOESM2_ESM.pdf-437-FOXO4_FOXO4	Rummagene Transcription Factor Associations 2026	1.0	null
CREB1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.858
EBF3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	33.1882
E2F4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	15.1171
SREBF1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	35.0781
PAX8	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.3573
TEAD4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	37.3768
OLIG2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	19.3231
TCF3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	29.2496
ETV5::FIGLA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	57.9224
THRA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	48.4615
FOSL1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	33.2172
FOSL2::JUNB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	33.1363
ZBTB12	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	16.6925
THRB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	108.972
IRF4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	26.8883
BATF::JUN	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.6495
TFAP4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	51.2473
JUN::JUNB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.1308
ZNF410	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	27.6808
NFIX	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	52.8229
SOX4	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	33.9445
GATA1::TAL1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	18.3443
KLF13	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	57.9073
TBX20	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	20.77
TTC1	CM4AI KOLF21J CRISPRi Gene Perturbation Atlas	-1.0	-2.35869
MGA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	22.7457
ZNF684	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	51.4283
TEAD1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	53.8163
BACH2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	36.9807
ZNF135	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	78.6582
FOSL1::JUND	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	36.2673
HOXC13	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	19.3147
HIC2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.6789
RFX3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	46.0193
MLX	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	20.6443
ZNF382	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	46.7564
TAL1::TCF3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	26.3864
IKZF1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	24.6345
ZNF24	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	40.6175
BHLHA15	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	26.4002
GFI1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	21.5108
MYOD1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	43.0875
DBP	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	19.7938
FIGLA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	16.3834
MYB	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	15.7996
chrX_72724424_G_A_b38	GTEx eQTL 2025	1.0	3.17244
CREB3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.8244
MAZ	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	76.4982
ZBTB26	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.8961
ETV1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	25.872
IRF2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	44.75
chrX_73088507_C_T_b38	GTEx eQTL 2025	1.0	1.984
TCF7L2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	50.0709
BATF	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.5626
ERF::FOXO1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	28.065
GSE114883_0_v_2_romidepsin_human	RummaGEO Drug Perturbation Signatures	-1.0	null
ETV2::FIGLA	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	39.2308
BATF3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	17.8041
MAFG::NFE2L1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	53.7867
PAX6	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	31.186
TEAD3	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	30.5383
PRPF4B	CM4AI KOLF21J CRISPRi Gene Perturbation Atlas	1.0	2.29125
ZBTB7C	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	29.176
ETV2::FOXI1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	27.6825
HEY2	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	18.1338
ZNF189	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	24.4363
ESR1	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	35.1912
ZBTB7A	JASPAR Predicted Human Transcription Factor Targets 2025	1.0	24.6436