A 20S multiprotein assembly of total mass about 1.2 MDa that activates dynein-based activity in vivo. A large structural component of the complex is an actin-like 40 nm filament composed of actin-related protein, to which other components attach. (Gene Ontology, GO_0005869)
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8 proteins localized to the dynactin complex cellular component from the COMPARTMENTS Curated Protein Localization Evidence Scores dataset.